Volume five continues the storyline we had in volume four, where Tanjiro and Inosuke are battling a family of spider demons. Father has gotten into the fight, and neither slayer can cut this demon with their katanas. Tanjiro and Inosuke get separated, and Tanjiro finds himself face to face with one of the Twelve Kizuki, the most powerful demons. Nezuko, in an effort to save Tanjiro, discovers a special power she has with her demon blood. Rui, the most powerful of the spider demons, becomes jealous of the familial bond between Tanjiro and Nezuko, and it angers him even more. Tanjiro’s katana breaks, while both of Inosuke’s katanas break battling Father. Giyu, the slayer from volume one, makes a triumphant comeback and saves Tanjiro from certain death. We also meet a new slayer, Shinobu, whose breathing methods take inspiration from insects, and who uses poisons to take down powerful demons.
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A mysterious, powerful warrior named Guts hunts down different demons and Apostles, powerful demons who have large segments of human population under their thrall. Guts inadvertently saves Puck, a tiny fairy, and gains a companion he doesn’t want. Puck tags along with Guts, at first because of a life-debt, and after that because he thinks he’ll see some amazing stuff in Guts’ company. Guts goes up against a powerful reptilian demon who is threatening a whole town and feasting off of its villagers. We find out later that Guts is also looking for more members of Godhand, a group that can be identified by the same brand he has on his neck.
Denji and his little devil dog, Pochita (who has a chainsaw coming out of his head), are swimming in a mountain of debt left by Denji’s dad. The two hunt devils to try and pay down the debt, but things go wrong for Denji on one such mission. Pochita saves Denji’s life by fusing with him, giving Denji the power to summon chainsaws from his head and arms! Now, the Public Safety Devil Hunters wants to employ this new, powerful Human-Devil hybrid, and Denji finds himself finally living a little comfortably. But, what now motivates him to keep the public safe? Is it money? Revenge? Boobs?
Illustrated by Posuka Demizu
Now that the world has been turned upside down, the Ratri clan desperately try to reestablish a similar government and farm system to put the world back to what it was. They plan on making all the farms like Lambda so none of the children they raise as food will have free will or rise up against them again. They prepare to make one last stand against Emma and her friends back where it all began: Grace Field House. Mom has now become Grandma, the head of the whole farm system, and she wields enormous power as the leader of the Sisters who work the farms. 
After clearing the house controlled by a demon with drums, Tanjiro and Zenitus come across another strange demon slayer, Inosuke, who likes to wear a warthog mask. Inosuke has an extremely hard head, both physically and metaphorically, and he can’t stand anyone doing anything better than him. Zenitsu reveals he knew Tanjiro was hiding a demon in his backpack, but trusts that Tanjiro needs to protect her. The growing group travel to a mountain to face an unknown demon threat, and they find other demon slayers caught up in a deadly web. They find a whole family of spider-type demons all over the mountain with many kinds of traps set for the slayers. The group, including hard-headed Inosuke, must work together to overcome the demons.
Somewhere in the world, someone is trying to raise the terrible demon lord, Gyumaoh, which made all the Yokai (demons) on Earth even more violent than before. A Buddhist monk, Genjyo Sanzo, and three warriors, Son Goku, Sha Gojyo, Cho Hakkai (all of whom have some Yokai blood in them), are tasked with stopping the revival and putting things back to normal. They travel through the world in a tiny dragon that can transform into a Jeep, and fight many battles along the way. The son of Gyumaoh tries to thwart the warriors, and he sends several of his henchmen to stop them or slow them down long enough for the revival to go off as planned.
Illustrated by Posuka Demizu
Norman's diabolical plan to eradicate the demons gets underway in this volume, despite Emma and Ray racing to the capital to try to stop him. First, Norman and his team blow up the bridges into the Demon Capital, so escaping is going to be tricky. Then, they invade the palace and pit the demon nobles up against each other along with their ally demon, Geelan, We also get all of the backstory on Geelan and how he came to be banished. Meanwhile, Emma and Ray navigate the demon capital trying to get to the castle to stop Norman before he goes too far. The Demon Queen, Legravalima, proves to be a hugely formidable foe, and her strength proves how she has managed to stay on top in a violent demon society. 
Musashi and his childhood friend, Kojiro, learn the legend of the Bushi - demon fighting warriors who use their katanas to fight even in the name of humanity. The only problem is, the two of them seem to be the only ones who think the demons are bad. Musashi hides his dream to become a Bushi and instead goes to school to become a miner, but he fashions his pickaxe to be a katana and continues to hone his skills in sword-fighting. On the day Musashi goes out to the mines, he discovers that his sword practice was well worth it, and that his dream of becoming a Bushi is much more obtainable than others made him believe.
Illustrated by Posuka Demizu
Emma and Ray have entered the playground of one of the most powerful demons in the world. They are there to renegotiate The Promise, the agreement between humans and demons that set the world up the way it is today. Norman is also prepared to march with a group of super-soldiers, who look strikingly like the incredibly strong Adam, and he is planning on taking his forces to the Imperial Capital of the demons. He sends Gilda and Don to find the “Evil-Blooded” demons, under the pretense that he wants to protect them from the war that is about to break out with all demon-kind. Don and Gilda know, however, that Norman wanted to get rid of the two nice demons, so they have to find a way to keep Sonja and Mujika safe. Emma finally cracks the riddle that will lead her to making a new promise, but she may have to sacrifice something much more than she’s willing in order to get it. 
Tanjiro and Nezuko help the demon doctors fight off two attacking demons, and we see more of the skills Tanjiro has learned in his training. Later, the two are off to their next assignment, but Tanjiro interrupts a desperate demon slayer trying to get a girl to marry him. This slayer, Zenitsu, is deathly afraid of demons and fighting, and asks Tanjiro to protect him on his mission. The two come across a house of demons where a few humans have been abducted. Outside are two siblings, and their brother is inside the house. Tanjiro and Zenitsu enter to try to save everyone. They get separated, and Zenitsu, despite his whining, reveals that he has extreme strength and speed, but only while he is sleeping. Tanjiro battles against the master of the house, who has drums on his body that shift the room and lob devastating attacks.
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